Afterbreakingoutofarehabilitationzoo,Frankie,asquirrelwithalltheangles,returnstohisforestHelearnsfromhisdimwittedbestfriend,Cody,aporcupineandstreetcon,thatwhilehewasinlockup,theentireforestwaspickedcleanbyTheACORNSInstituteAlternativeCosmeticOilsforRegenerativeNaturalSkincareNoacornwasleftbehindWithwinterjustaroundthecorner,it'sapotentiallydevastatingblowtoallforestcrittersSo,Frankiehasnochoicebuttobustintothebrandnew,stateoftheart,rodentproofACORNSProcessingPlantandtakebackalloftheacornsThoughaseeminglyimpossibleheist,successwouldnotonlysavethelivesofhisforestfriends,butitwouldalsogiveFrankieachancetowinbacktheloveofhislife,Lola,alocalshowsquirrelSadly,LolahasbeenburntbytoomanyofFrankie'sfalsepromisesinthepastWiththesupportofFrankie'soldmentor,MrBellwood,awealthybadger,FrankieandCodyassembleateamoftopnotchcrackersfromallovertheforest,includingabatnamedFlyBoy,atopnotchmunitionsexpert;Raitch,acomputerhackingalbinosnake,whoneverallowsherlackofappendagestogetintheway;Edsyatinyfamilyfrog,whoseabilitytofreezehimselfsolidfordaysonendwithoutharmisinvaluable;agermophobicFrenchratnamedLiam,whocancollapsehisskeletontogetintoanyspace;ChinesefatherdaughterpangolinsnamedPangoandLin,thebestdiggersintheforest;andBellwood'shedgehogbutler,OscarwhoknowshowtogethishandsonanythingthecrewneedsWithwinterknockingattheirdoor,FrankieandhiscrewhaveonlythreedaystoprepareandexecutethisunprecedentedheistWhentheylearnthattheprocessingoftheirforest'sacornshasbeenpushedupaday,Lola'srefusaltoparticipateleavesFrankieandhiscrewonecrittershortWitheverythingontheline,FrankieandhiscrewofAnimalCrackerzsetouttopulloffthebiggestheistinanimalhistory
